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C. ramosus Chaudhuri et al. 1992 immatures

Larva:  a small to medium plumosus-type (length 8-13.2 mm).   Anterior pair of ventral tubules shorter than posterior pair.  Anal tubules 310 (290-330) µm long.  Gula and frontoclypeus apparently not darkened.
Mentum with c2 teeth well separated and c1 broad (type IB), 4th lateral reduced (type II).
Ventromentum about 3.9 times longer than its depth; figure indicates numerous striae.  Pecten epipharyngis with 20 teeth (type B) - other populations have 12-16 teeth.  Premandible shown with two equal teeth, but normally the inner tooth is 2-3 times wider than the outer tooth (perhaps Ty. D).
Antenna with basal segment about 2.25 times longer than wide, RO less than 1/3 up from the base; AR 1.69 (1.53-2.10); ratio of antennal segments 23.1 : 6.7 : 2.2 : 3.1 : 1.6.
Mandible figured as type IIB; no information on number of furrows on outer surface; or number of taeniae in PMa; MTR perhaps about 0.33.


Illustration of larval parts of C. ramosus from Chaudhuri et al.1992

Pupa: Exuvia grey.  Length of male 6.69 (6.67-7.64) mm, of female 7.27 (6.52-8.86).  Frontal tubercles about 0.09 mm long and 0.06 mm wide, with the subapical seta about 0.05 mm in length.  Antennal sheath of female about 540 (540-560) µm long.  Wing sheath 1.54 (1.42-1.72) mm in length, respiratory base elliptical and about 0.15 mm wide.  PSA caudolateral on segment IV-VI; PSB basolateral on segment I and caudolateral on segment II.  Hook row on segment II comprised of about 46-58 hooklets.  Shagreen pattern as shown in figure.  Caudolateral spurs of segment VIII with 3 spines.

Cytology: Four polytene chromosomes with the pseudothummi-cytocomplex combination BF, CD, AE, G.
Although later specimens attributed to C. ramosus (e.g. Laviad-Shirit et al., 2020; Sela et al. 2021) are actually C. flaviplumus Ty. B, the original description by Nath & Godbole (1997) is C. ramosus.  In their map, chromosome I is comprised of arms D and C, chromosome II of arms F and B; chromosome III of arms A and E, and chromosome IV is arm G.  There are two nucleoli, although the one in arm C is development stage specific.  The other is in arm B at 11B.  Two BRs, about equally distant from each end, are on arm G.  Band identifications as in Nath & Godbole (1997):
Nucleolus in arm B (at 11B)(as in map of Nath & Godbole 1997).  Two BRs, about equally distant from each end, are on arm G.  No reported polymorphism.
Some points can be made by reference to the map of Nath & Godbole (1997):
ramA1:    1 - 2c, 10 - 12, 3- 2d, 9 - 4, 13 - 19                as circumdatus A2, holomelas, etc.
ramB1:    Characteristic bands at 10F.  Nucleolus at 11B
ramC1:    Developmentally stage specific nucleolus at 6B.            as C2 of incertipenis?
ramD1:   
ramD2:    Simple inversion towards end of armramE1:    1a-i, 5 - 10b, 3e - 2, 4 - 3f, 10c - 13            i.e Inv2-5 from aprilinus, as incertipenis
ramF1:    1 - 2a, 10 - 3d, 14c - 11, 2b - 3c, 14d - 23.
ram G1:    BRs at 18C and 19C


Chromosomes of C. ramosus, courtesy of B. B. Nath. N - nucleolus; BR - Balbiani ring

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Modified: 26 November 2025
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