A listing of Chironomidae reported from California

Version of November 1999

The present edition contains 91 genera and 245 species. 19 of the latter (species name underlined) to my knowledge have not previously been reported from California.

The taxa are arranged alphabetically, as that is the system least subject to change over time and differences in opinion.

A second table column on the right gives references for California occurrence of the respective species on the same line. Unfortunately, some browsers may be unable to perfectly reproduce the two-column layout of my source file. If you have any problems, you are welcome to email me for an orderly copy.

Wherever there is no entry in the right-hand column, California has already been specifically included in that particular species' distribution by the latest Nearctic catalog (Oliver, Dillon & Cranston 1990, Oliver & Dillon 1994; in the following referred to as "NACAT"). For species listed by NACAT as "widespread", I give at least one source for actual records from California. An entry like "Smith (1975), not NACAT" means that in contrast to the reference cited, distribution according to NACAT does not include California or any neighboring state. Unpublished personal observations are shown as "MS".

Every time the list will be updated, alterations will be signalled and dated (month/year) - example: New 5/2000 - allowing you to quickly find changes since the latest version you have seen.

Whenever you notice an error, publish a new California record, or have any other comment, please don't hesitate to contact me. And if you download and work with this file, kindly do not distribute any copies without my consent if you have made alterations I don't know of. Thank you for your understanding!

Taxa References for California
CHIRONOMINAE
Chironomini
Apedilum Townes, 1945
elachistum Townes, 1945
subcinctum Townes, 1945
Chernovskiia Sæther, 1977
orbicus (Townes, 1945)
Chironomus Meigen, 1803
anonymus Williston, 1896
anthracinus Zetterstedt, 1860
atrella (Townes, 1945)
calligraphus Goeldi, 1905 Spies & Reiss (1996), not NACAT
cucini Webb, 1969
decorus Johannsen, 1905 Townes (1945), not NACAT
frommeri Sublette & Sublette, 1971
longipes Staeger, 1839 Sublette & Sublette (1965), not NACAT
maturus Johannsen, 1908
mendax Storå, 1936
plumosus (Linné, 1758) Sublette (1960)
riparius Meigen, 1804 Sublette (1960)
staegeri Lundbeck, 1898
stigmaterus Say, 1823
tuxis Curran, 1930 Whitsel et al. (1963), not NACAT
utahensis Malloch, 1915
whitseli Sublette & Sublette, 1974
Cladopelma Kieffer, 1921
amachaerus (Townes, 1945)
edwardsi (Kruseman, 1933)
forcipis (Rempel, 1939) Lothrop & Mulla (1995), not NACAT
viridulum (Linné, 1767)
Cryptochironomus Kieffer, 1918
curryi Mason, 1986 Darby (1962) according to Mason (1986)
digitatus (Malloch, 1915) Townes (1945), not NACAT
fulvus (Johannsen, 1905)
ponderosus (Sublette, 1964) MS, not NACAT
Cryptotendipes Beck & Beck, 1969
ariel (Sublette, 1960)
darbyi (Sublette, 1960)
Dicrotendipes Kieffer, 1913
californicus (Johannsen, 1905)
crypticus Epler, 1987
fumidus (Johannsen, 1905) Epler (1987), not NACAT
modestus (Say, 1823) Sublette (1960), Epler (1988)
nervosus (Staeger, 1839)
tritomus (Kieffer, 1916) Epler (1987), not NACAT
Endochironomus Kieffer, 1918
nigricans (Johannsen, 1905)
Endotribelos Grodhaus, 1987
hesperium (Sublette, 1960)
Glyptotendipes Kieffer, 1913
barbipes (Staeger, 1839) Grodhaus (1967), not NACAT
lobiferus (Say, 1823)
Goeldichironomus Fittkau, 1965
amazonicus (Fittkau, 1968) Sublette & Mulla (1991), not NACAT
holoprasinus (Goeldi, 1905)
Harnischia Kieffer, 1921
curtilamellata (Malloch, 1915) Sublette (1960)
Kiefferulus Goetghebuer, 1922
modocensis (Sublette, 1960)
Lauterborniella Thienemann & Bause, 1913
agrayloides (Kieffer, 1911)
Microchironomus Kieffer, 1918
nigrovittatus (Malloch, 1915)
Microtendipes Kieffer, 1915
caducus Townes, 1945
pedellus (De Geer, 1776)
Nilothauma Kieffer, 1920
babiyi (Rempel, 1937) MS, not NACAT
mirabile (Townes, 1945) MS, not NACAT
Parachironomus Lenz, 1921
abortivus (Malloch, 1915) MS, not NACAT
chaetoalus (Sublette, 1960)
directus (Dendy & Sublette, 1959) Lothrop & Mulla (1995), not NACAT
frequens (Johannsen, 1905) Sublette (1960)
hirtalatus (Beck & Beck, 1964) Lothrop & Mulla (1995), not NACAT
tenuicaudatus (Malloch, 1915) Sublette (1960), Frommer & Sublette (1971)
Paracladopelma Harnisch, 1923
alphaeus (Sublette, 1960)
Paratendipes Kieffer, 1911
albimanus (Meigen, 1818)
fuscitibia Sublette, 1960
subaequalis (Malloch, 1915)
thermophilus Townes, 1945
Phaenopsectra Kieffer, 1921
dyari (Townes, 1945)
flavipes (Meigen, 1818)
mortensoni Grodhaus, 1987
pilicellata Grodhaus, 1976
profusa (Townes, 1945)
Polypedilum Kieffer, 1912
albicorne (Meigen, 1838)
albinodus Townes, 1945
angustum Townes, 1945
apicatum Townes, 1945
artifer (Curran, 1930)
aviceps Townes, 1945
californicum Sublette, 1960
cinctum Townes, 1945
cultellatum Goetghebuer, 1932 Oyewo & Sæther (1998), not NACAT
digitifer Townes, 1945 Soponis & Simpson (1992)
halterale (Coquillett, 1901) Sublette (1960)
illinoense (Malloch, 1915)
isocerus Townes, 1945
labeculosum (Mitchell, 1908)
laetum (Meigen, 1818)
obelos Sublette & Sasa, 1994 MS, not NACAT
ophioides Townes, 1945
parvum Townes, 1945
pedatum Townes, 1945
scalaenum (Schrank, 1803) Sublette (1960)
sulaceps Townes, 1945
tritum (Walker, 1856)
Robackia Sæther, 1977
demeijerei (Kruseman, 1933) Leland & Fend (1998), not NACAT
Sergentia Kieffer, 1922
albescens (Townes, 1945)
Stenochironomus Kieffer, 1919
colei (Malloch, 1919)
fuscipatellus Borkent, 1984
totifuscus Sublette, 1960
Stictochironomus Kieffer, 1919
naevus (Mitchell, 1908)
quagga (Townes, 1945)
Synendotendipes Grodhaus, 1987
luski Grodhaus, 1987
Tribelos Townes, 1945
jucundum (Walker, 1848)
subatrum Grodhaus, 1987
subletteorum Grodhaus, 1987
Pseudochironomini
Pseudochironomus Malloch, 1915
richardsoni Malloch, 1915
Tanytarsini
Caladomyia Säwedal, 1981
pistra Sublette & Sasa, 1994 Lothrop & Mulla (1995), not NACAT
Cladotanytarsus Kieffer, 1921
viridiventris (Malloch, 1915)
Micropsectra Kieffer, 1909
nigripila (Johannsen, 1905)
polita (Malloch, 1915)
Paratanytarsus Thienemann & Bause, 1913
grimmii (Schneider, 1885) MS, not NACAT
Tanytarsus van der Wulp, 1874
angulatus Kawai, 1991 Spies (1998a)
challeti Spies, 1998 Spies (1998a)
dendyi Sublette, 1964 Ali & Mulla (1976), not NACAT
hastatus Sublette & Sasa, 1994 Spies & Reiss (1996)
limneticus Sublette, 1964 Lothrop & Mulla (1995), not NACAT
mendax Kieffer, 1925 Spies (1998b), not NACAT
neoflavellus Malloch, 1915 Anderson et al. (1964), not NACAT
pelsuei Spies, 1998 Spies (1998a)
   
DIAMESINAE
Diamesini
Diamesa Meigen, 1835
chorea Lundbeck, 1898
davisi Edwards, 1933 Hansen & Cook (1976), not NACAT
heteropus (Coquillett, 1905) Hansen & Cook (1976)
japonica Tokunaga, 1936
sonorae Willassen, 1985
spinacies Sæther, 1969
Pseudodiamesa Goetghebuer, 1939
branickii (Nowicki, 1873)
Sympotthastia Pagast, 1947
diastena (Sublette, 1964)
   
ORTHOCLADIINAE
Brillia Kieffer, 1913
parva (Johannsen, 1934) Frommer & Sublette (1971), not NACAT
retifinis Sæther, 1969
Cardiocladius Kieffer, 1912
platypus (Coquillett, 1902)
Chasmatonotus Loew 1864
hyalinus Coquillett, 1905
maculipennis Rempel, 1937
univittatus Coquillett, 1900
Clunio Haliday, 1855
californiensis Hashimoto, 1974
Cricotopus van der Wulp, 1874
annulator Goetghebuer, 1927
bicinctus (Meigen, 1818) Darby (1962)
blinni Sublette, 1998 Sublette et al. (1998)
furtivus Sublette & Sublette, 1971
fuscatus Wirth, 1957
globistylus Roback, 1957 Sublette et al. (1998), not NACAT
herrmanni Sublette, 1998 Sublette et al. (1998)
infuscatus (Malloch, 1915) Sublette et al. (1998), not NACAT
nostocicola Wirth, 1957
obscurifuscus Sublette & Sublette, 1971
ornatus (Meigen, 1818) MS, not NACAT
parafuscatus Sublette & Sublette, 1971
subletteorum Spies, 1999 Spies (1999)
sylvestris (Fabricius, 1794) Darby (1962)
trifascia Edwards, 1929 Sublette et al. (1998), not NACAT
Doithrix Sæther & Sublette, 1983
barberi Sæther & Sublette, 1983
ensifer Sæther & Sublette, 1983
Eretmoptera Kellogg, 1900
browni Kellogg, 1900
Eukiefferiella Thienemann, 1926
claripennis (Lundbeck, 1898) MS, not NACAT
coerulescens (Kieffer, 1926) MS, not NACAT
devonica (Edwards, 1929)
ilkleyensis (Edwards, 1929) MS, not NACAT
Euryhapsis Oliver, 1981
annuliventris (Malloch, 1915)
illoba Oliver, 1981
Georthocladius Strenzke, 1941
platystylus Sæther & Sublette, 1983
wirthi Sæther & Sublette, 1983
Heterotrissocladius Spärck, 1923
oliveri Sæther, 1975
Hydrobaenus Fries, 1830
pilipes (Malloch, 1915)
Limnophyes Eaton, 1875
asquamatus Andersen, 1937
natalensis (Kieffer, 1914) MS, not NACAT
Metriocnemus van der Wulp, 1874
edwardsi Jones, 1916
Nanocladius Kieffer, 1913
anderseni Sæther, 1977 MS, not NACAT
Orthocladius van der Wulp, 1874
appersoni Soponis, 1977
carlatus (Roback, 1957)
dubitatus Johannsen, 1942 Sublette et al. (1998), not NACAT
frigidus (Zetterstedt, 1838) Sublette et al. (1998), not NACAT
hellenthali Soponis, 1977
lignicola Kieffer, 1915 Soponis (1977), not NACAT
mallochi Kieffer, 1919
obumbratus (Johannsen, 1905)
rubicundus (Meigen, 1818) Soponis (1977), not NACAT
subletti Soponis, 1977
Parachaetocladius Wülker, 1959
imberbus Sæther & Sublette, 1983
Parakiefferiella Thienemann, 1936
subaterrima (Malloch, 1915) Sublette et al. (1998), not NACAT
Parametriocnemus Goetghebuer, 1932
lundbeckii (Johannsen, 1905)
Paraphaenocladius Spärck & Thienemann, 1924
exagitans (Johannsen, 1905) Sæther & Wang (1995), not NACAT
innasus Sæther & Wang, 1995 Sæther & Wang (1995)
Paratrichocladius Santos Abreu, 1918
rufiventris (Meigen, 1830) MS, not NACAT
Psectrocladius Kieffer, 1906
barbimanus (Edwards, 1929) MS, not NACAT
spinifer (Johannsen, 1928)
vernalis Malloch, 1915 Lothrop & Mulla (1995), not NACAT
Pseudorthocladius Goetghebuer, 1932
dumicaudus Sæther, 1969
uniserratus Sæther & Sublette, 1983
Pseudosmittia Edwards, 1932
forcipata (Goetghebuer, 1921) MS, not NACAT
nanseni Kieffer, 1926
Symbiocladius Kieffer, 1925
equitans (Claassen, 1922)
Tethymyia Wirth, 1949
aptena Wirth, 1949
Thalassosmittia Strenzke & Remmert, 1957
clavicornis (Saunders, 1928)
marina (Saunders, 1928)
pacifica (Saunders, 1928)
Tvetenia Kieffer, 1922
vitracies (Sæther, 1969) Sublette et al. (1998), not NACAT
   
PODONOMINAE
Boreochlini
Boreochlus Edwards, 1938
persimilis (Johannsen, 1926)
sinuaticornis Brundin, 1966
Podonomini
Parochlus Enderlein, 1912
kiefferi (Garrett, 1925)
   
TANYPODINAE
Coelotanypodini
Clinotanypus Kieffer, 1913
pinguis (Loew, 1861) Roback (1976)
Macropelopiini
Alotanypus Roback, 1971
venustus (Coquillett, 1902)
Apsectrotanypus Fittkau, 1962
algens (Coquillett, 1902) Wirth & Stone (1956), not NACAT
Brundiniella Roback, 1978
eumorpha (Sublette, 1964)
Derotanypus Roback, 1971
aclines (Sublette, 1964)
Psectrotanypus Kieffer, 1909
dyari (Coquillett, 1902)
Radotanypus Fittkau & Murray, 1985
florens (Johannsen, 1908)
submarginellus (Sublette, 1964)
Natarsiini
Natarsia Fittkau, 1962
miripes (Coquillett, 1905)
Pentaneurini
Ablabesmyia Johannsen, 1905
annulata (Say, 1823)
aspera (Roback, 1959)
cinctipes (Johannsen, 1946)
mallochi (Walley, 1925)
monilis (Linné, 1758) Sublette (1964), Roback (1971)
peleensis (Walley, 1926) Sublette (1964)
Conchapelopia Fittkau, 1957
pallens (Coquillett, 1902)
Hayesomyia Murray & Fittkau, 1985
senata (Walley, 1925)
Labrundinia Fittkau, 1962
maculata Roback, 1971
pilosella (Loew, 1866) Sublette (1964), Roback (1987)
Larsia Fittkau, 1962
decolorata (Malloch, 1915) MS, not NACAT
lyra (Sublette, 1964)
marginella (Malloch, 1915) Wirth & Stone (1956) , not NACAT
planensis (Johannsen, 1946)
sequoiaensis (Sublette, 1964)
Meropelopia Roback, 1971
flavifrons (Johannsen, 1905)
Monopelopia Fittkau, 1962
tenuicalcar (Kieffer, 1918)
Nilotanypus Kieffer, 1923
fimbriatus (Walker, 1848) Roback (1986)
Paramerina Fittkau, 1962
smithae (Sublette, 1964)
Pentaneura Philippi, 1865
inconspicua (Malloch, 1915)
inyoensis Sublette, 1964
Thienemannimyia Fittkau, 1957
barberi (Coquillett, 1902)
fusciceps (Edwards, 1929)
Xenopelopia Fittkau, 1962
tincta Roback, 1971
Zavrelimyia Fittkau, 1962
sinuosa (Coquillett, 1905)
thryptica (Sublette, 1964)
Procladiini
Procladius Skuse, 1889
barbatulus Sublette, 1964
bellus (Loew, 1866)
culiciformis (Linné, 1767)
denticulatus Sublette, 1964
freemani Sublette, 1964
sublettei Roback, 1971
Tanypodini
Tanypus Meigen, 1803
carinatus Sublette, 1964
grodhausi Sublette, 1964
imperialis Sublette, 1964
neopunctipennis Sublette, 1964 Lothrop & Mulla (1995) , not NACAT
nubifer Coquillett, 1905
parastellatus Sublette, 1964
punctipennis Meigen, 1818
stellatus Coquillett, 1902 Sublette (1964), Roback (1971), not NACAT
   
TELMATOGETONINAE
Telmatogeton Schiner, 1866
alaskense Coquillett, 1900
macswaini Wirth, 1949
spinosum (Hashimoto, 1973)
trilobatum (Kieffer, 1911)

REFERENCES

Martin Spies
Schraemelstr. 151
D-81247 Muenchen
Germany

spies@zi.biologie.uni-muenchen.de