Welcome to the Chironomid Exchange Forum! Use this resource to discuss midge matters with the world-wide community of researchers, and to stay up-to-date on important data, e.g. in standard reference publications.
Please report to moderators any spammers or attempts to use this forum for purposes other than the exchange of scientific information related to the science of Chironomidae or entomology. Thank you!
Ethan and Martin - Moderators

Main Menu

Recent posts

Various topics / Re: GENERAL INFORMATION
Last post by Martin Spies - January 22, 2020, 12:32:00 PM
Dear all,

I would like to inform you that Dr. Tadashi Kobayashi passed away on 10 January, at age 88 years.

Masaru Yamamoto
Recently published / Re: Orthocladiinae - new taxa
Last post by Martin Spies - January 15, 2020, 01:50:51 PM

Donato, M., Siri, A. (2019) Phylogeny and new species of the genus Lipurometriocnemus Sæther (Diptera: Chironomidae). Arthropod Systs & Phyl. 77(2): 339-349 + online suppl. material. [first publd online 17.ix.2019]

Krasheninnikov, A. B. (2019) Zelentsoviella jugorica gen. nov, sp. nov., a new Orthocladiinae from Russian Arctic (Diptera, Chironomidae). Zootaxa 4585(3): 581-590. [publd 15.iv.2019]

Makarchenko, E. A., Makarchenko, M. A. (2019) Obzor khironomid roda Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) rossiiskogo Dal'nego Vostoka. (Review of the genus Gymnometriocnemus Edwards (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East). Evraziat. Entomol. Zh. (= Euroas. Entomol. J.) 18(5): 333-340. [in Russian, with English abstract]

Makarchenko, E. A., Makarchenko, M. A., Pozdeev, I. V., Yavorskaya, N. M. (2019) Novye dannye po taksonomii khironomid roda Eukiefferiella Thienemann (Diptera, Chironomidae, Orthocladiinae) rossiiskogo Dal'nego Vostoka. (New data on taxonomy of Eukiefferiella Thienemann (Diptera, Chironomidae, Orthocladiinae) from the Russian Far East). Evraziat. Entomol. Zh. (= Euroas. Entomol. J.) 18(1): 47-59. [in Russian, with English abstract]

Makarchenko, E. A., Makarchenko, M. A., Semenchenko, A. A. (2019a) Towards the taxonomy of Corynoneura Winnertz (Diptera: Chironomidae: Orthocladiinae) from the Russian Far East and eastern Siberia. Zootaxa 4612(2): 221-236. [publd 29.v.2019]

Moubayed, J., Kettani, K. (2019) Rheocricotopus (Rheocricotopus) rifensis sp. n., a new rheobiontic species from waterfalls located in the Moroccan Rif [Diptera, Chironomidae, Orthocladiinae]. Ephemera 20(2): 87-97. ["2018"; in English, with French abstract]

Moubayed, J., Langton, P. H. (2019) Chaetocladius berythensis sp. n., C. callauensis sp. n., C. guardiolei sp. n. and C. parerai sp. n., four relict species inhabiting glacial springs and streams in eastern Pyrenees and Lebanon (Diptera: Chironomidae). CHIRONOMUS J. Chironomidae Res. 32: 42-59. [publd online (8.xi.2019)]

Moubayed, J., Tissot, B. (2019) Smittia remoraya sp. n., a new semiterrestrial species inhabiting alkaline wet sedge meadows in north eastern France [Diptera, Chironomidae, Orthocladiinae]. Ephemera 20(2): 77-85. ["2018"; in English, with French abstract]

Moubayed-Breil, J., Ashe, P. (2019) Rheocricotopus (s. str.) costai sp. n. and R. (s. str.) pyrenaeus sp. n., two relict species from glacial rheocrenes and streams in Corsica and the eastern Pyrenees (Diptera: Chironomidae, Orthocladiinae). Trakya Univ. J. nat. Scis: 12 pp. [first publd online (13.iii.2019), with pagin. in issue 20(1): 35-46 (iv.2019)]

Moubayed-Breil, J., Bitušík, P. (2019) Taxonomic notes on the genus Chaetocladius (laminatus-group) (Diptera: Chironomidae, Orthocladiinae) II. Descriptions of C. bitusiki sp. n. and C. mantetensis sp. n., two relict species inhabiting cold alpine springs and streams. Biologia 74(11): 1489-1500. [publd +30.xi.2019; the version publd online earlier did not meet the nomenclatural requirements]

Moubayed-Breil, J., Dominici, J.-M. (2019) Clunio boudouresquei sp. n. and Thalassosmittia ballestai sp. n., two Tyrrhenian marine species occurring in Scandola Nature Reserve, West Corsica (Diptera: Chironomidae). CHIRONOMUS J. Chironomidae Res. 32: 4-24. [publd online (4.x.2019)]

Moubayed-Breil, J., Mary, N. (2019) Eukiefferiella coconina sp. n., an afrotropical element occurring in eurythermal lotic habitats of Mayotte Island, France [Diptera, Chironomidae, Orthocladiinae]. Ephemera 20(1): 3-17. ["2018"; in English, with French abstract]

Yan, Ch., Liu, T., Cao, W., Zhang, X., Liu, W. (2019) Review of Epoicocladius Šulc et Zavrel from China (Diptera, Chironomidae). Zootaxa 4590(2): 289-295.
Various topics / POSITION announcements
Last post by Martin Spies - January 11, 2020, 02:17:27 PM
Dear colleagues,

The following is posted here upon request by the colleagues hoping to find someone to hire.


Chironomidae Taxonomist
EcoAnalysts, Inc. Moscow, Idaho

EcoAnalysts is currently searching for an individual who enjoys chironomidae taxonomy and who wants to be part of a successful team of aquatic scientists. If you wish to use your knowledge and experience to help people make highly informed decisions regarding the condition and stewardship of our natural resources, then we encourage you to explore the information below and throughout our entire website at

We are currently seeking qualified applicants to process chironomidae samples.  Preference will be given to those who demonstrate experience and knowledge of identifying chironomidae.  Duties include identification of samples in a production environment and data summarization for reporting to clients in addition to related laboratory needs.



The major responsibility of the Chironomidae Taxonomist is the identification of chironomidae samples; therefore, demonstrated experience with chironomidae taxonomy is required.
Knowledge of appropriate sample handling and preparation techniques is required.
Proficient reading and writing skills and the ability to follow protocols.  Experience using literature to make ID's.
Effective communication skills both verbal and in writing.
Experience with North American Chironomidae taxa from a variety of habitats.
Accuracy and speed is imperative due to fixed project budgets.
Excellent work ethic, ability to meet deadlines, willingness to go the "extra mile", and ability to work well in small groups and independently.
Minimum education requirement is a BS in a related scientific field, but preference will be given to higher education levels. Preference will also be given to applicants with at least two years of identification experience in a production taxonomy setting.

Salary range and benefits are dependent upon experience, qualifications and job performance. Benefits include flex time, vacation/sick time, a health insurance plan (medical, vision and dental) and an employer-matched investment fund.

Please send or email 1) letter of application, 2) complete resume, and 3) three professional references.


Anticipated start date
Positions available immediately

Megan Payne
Laboratory Manager
EcoAnalysts, Inc.
1420 S. Blaine Street, Ste 14
Moscow, Idaho 83843
Molecules / Re: Recent publications on mol...
Last post by Martin Spies - January 01, 2020, 03:14:42 PM

Czechowski, P., Stevens, M. I., Madden, C., Weinstein, P. (2019) Steps towards a more efficient use of chironomids as bioindicators for freshwater bioassessment: Exploiting eDNA and other genetic tools. Ecol. Indicators 110: 11 pp. + online suppl. material. [first publd online (20.xi.2019)]

Gajski, G., Žegura, B., Ladeira, C., Pourrut, B., Del Bo', C., Novak, M., Sramkova, M., Milic, M., Gutzkow, K. B., Costa, S., Dusinska, M., Brunborg, G., Collins, A. (2019) The comet assay in animal models: From bugs to whales – (Part 1 Invertebrates). Mutation Res. / Revs Mutation Res. 779: 82-113.

McCulloch, G. A., Purcell, M. F., Harms, N. E., Grodowitz, M. J., Zhang, J., Hong, S.-H., Walter, G. H. (2019) Molecular screening of herbivorous flies collected from Hydrilla verticillata across China and Korea – setting up hypotheses for further exploratory surveys and tests. Biol. Control 138(paper 104051): 6 pp. + online suppl. material.

Morinière, J., Balke, M., Doczkal, D., Geiger, M. F., Hardulak, L. A., Haszprunar, G., Hausmann, A., Hendrich, L., Regalado, L., Rulik, B., Schmidt, S., Wägele, J., Hebert, P. D. (2019) A DNA barcode library for 5,200 German flies and midges (Insecta: Diptera) and its implications for metabarcoding-based biomonitoring. Mol Ecol Resour.: 29 pp. + online suppl. material. [first publd online (12.iv.2019);]

Weigand, H., Beermann, A. J., Čiampor, F., Costa, F. O., Csabai, Z. et al. [41 authors in total, incl. several chironomid researchers] (2019) DNA barcode reference libraries for the monitoring of aquatic biota in Europe: Gap-analysis and recommendations for future work. Sci. Total Envir. 678: 499-524 + online suppl. material.
Last post by Martin Spies - December 27, 2019, 11:21:21 AM
For an update, please see

Cornette, R. (2019) The 21st International Symposium on Chironomidae. CHIRONOMUS J. Chironomidae Res. 32: 84-85. [publd online (26.xii.2019)]

This paper can be downloaded from here:
Recently published / Re: Chironominae - new taxa
Last post by Martin Spies - December 23, 2019, 01:56:00 PM

Andersen, T., Pinho, L. C. (2019) A new species of Fissimentum Cranston & Nolte, 1996 from the state of São Paulo, Brazil (Diptera: Chironomidae, Chironominae). Biotemas 32(4, Dec.): 45-49. [in English, with Portuguese abstract] [This online-only publication did not meet the corresponding nomenclature requirements (e.g., see ICZN 2012 in Zootaxa 3450, esp. concerning Code Article 8.5). Consequently, the species name it proposes was not made available. The same applies to names in Lipurometriocnemus proposed in the same journal by Andersen et al. (2016).]

Cranston, P. S. (2019a) Paraskusella gen. nov., a new Afro-Australian genus in the tribe Chironomini (Diptera: Chironomidae). Austral Ent. 58(2): 268-281.

Cranston, P. S. (2019b) Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific Region. Zootaxa 4646(3): 461-500. [publd 25.vii.2019]

Fusari, L. M., Dantas, G. P. S., Hamada, N., Andrade-Ouza[sic!], V., Lima, K. M., Silva, J. G. (2019) Erratum. LÍVIA MARIA FUSARI, GALILEU P.S. DANTAS, NEUSA HAMADA, VANDERLY ANDRADE-OUZA[sic!], KÁTIA M. LIMA & JANISETE G. SILVA (2018) Not endemic after all: Imparipecten Freeman, 1961 (Diptera: Chironomidae) described from the Neotropical Region . Zootaxa, 4532 (3): 396–406 .[sic!] - Zootaxa 4712(4): 600. [publd 23.xii.2019]

Lin, X.-L., Yu, H.-J., Zhang, R.-L., Wang, X.-H. (2019) Polypedilum (Cerobregma) heberti sp. n. (Diptera: Chironomidae) from Gaoligong Mountains, Yunnan, China. Zootaxa 4571(2): 255-262. [publd 26.iii.2019; for Erratum, see Lin et al. (2020)]

Mukherjee, T., Pal, G., Hazra, N. (2019) On a new species of the genus Dicrotendipes Kieffer, 1913 from West Bengal, India with cladistic analysis and key to the Oriental species (Diptera: Chironomidae). Oriental Insects 54(2): 216-240. [first publd online 28.v.2019, with issue pagin. 2.iv.2020]

Pinho, L. C., Dantas, G. P. S., Hamada, N. (2019) Some notes on taxonomy and distribution of Neotropical Aedokritus Roback and Axarus Roback (Diptera: Chironomidae: Chironominae). Zootaxa 4668(4): 535-542. [publd 12.ix.2019]

Shimabukuro, E. M., Trivinho-Strixino, S., Lamas, C. J. E. (2019) New Polypedilum Kieffer (Diptera: Chironomidae) from mountains of the Atlantic Forest, Brazil. Zootaxa 4612(4): 518-532. [publd 31.v.2019]

Tang, H., Cranston, P. S. (2019) A new tribe in the Chironominae (Diptera: Chironomidae) validated by first immature stages of Xiaomyia Sæther & Wang and a phylogenetic review. Raffles Bull. Zool. 67: 684-693. [publd 4.xii.2019]
Established earlier / Re: New publications on variou...
Last post by Martin Spies - December 20, 2019, 04:05:16 PM

Borkent, A. (2ß19) The Lower Cretaceous male of Lebanoculicoides daheri – belonging to the earliest lineage of biting midges (Diptera: Ceratopogonidae). Can. Ent.: 13 pp. [first publ. online (21.iii.2019); doi:10.4039/tce.2019.4]

Cranston, P. S. (2019) An enigmatic Chironomidae (Diptera): first larval description for Nandeva Wiedenbrug, Reiss and Fittkau and evidence for phylogenetic position in Tanytarsini. CHIRONOMUS J. Chironomidae Res. 32: 25-32. [publd online (3.xi.2019)]

Donato, M., Baccaro, L., Siri, A. (2019) Male description of the phytotelma-dwelling species Polypedilum parthenogeneticum (Chironomidae: Chironominae). CHIRONOMUS J. Chironomidae Res. 32: 68-72. [publd online (11.x.2019)]

Evenhuis, N. L., Pape, T. (2019) Nomenclatural studies toward a world list of Diptera genus-group names. Part VII: Johann Wilhelm Meigen. Zootaxa 4703(1): 001–193. [publd 5.xii.2019]

Fernandes, I. da S., Nicacio, G., Rodrigues, G. G., Silva, F. L. da (2019) Freshwater sponge-dwelling Chironomidae (Insecta, Diptera) in northeastern Brazil. Biotemas 32(1, March): 39-48.

Han, W., Tang. H. (2019) Phylogeny of marine Ainuyusurika tuberculata (Tokunaga) (Diptera: Chironomidae: Chironominae), with description of the immature stages. Zootaxa 4695 (2): 131–147. [publd 7.xi.2019]

Laug, A., Hamerlík, L., Anslan, S., Engels, S., Turner, F., Wang, J., Schwalb, A. (2019) Acricotopus indet. morphotype incurvatus: Description and genetics of a new Orthocladiinae (Diptera: Chironomidae) larval morphotype from the Tibetan Plateau. Zootaxa 4656(3): 535-544.

Lin, X.-L., Wang, X.-H. (2019) Thienemannimyia (Thienemannimyia) sinogalbina nom. n., a replacement name for Thienemannimyia galbina Cheng & Wang, 2009. CHIRONOMUS J. Chironomidae Res. 32: 73-74. [publd online (16.x.2019)]

Makarchenko, E. A., Makarchenko, M. A., Semenchenko, A. A. (2019b) Morphological description and DNA barcoding of Hydrobaenus laticaudus Sæther, 1976 (Diptera: Chironomidae: Orthocladiinae) from Amur River basin (Russian Far East). Zootaxa 4674(2): 225-234. [publd 24.ix.2019]

Namayandeh, A., Beresford, D. V. (2019) Chironomidae of Fosheim Peninsula, Ellesmere Island: first female description of Chaetocladius (Chaetocladius) glacialis (Lundström, 1915) (Diptera: Chironomidae: Orthocladiinae) and new faunistic records from the Nunavut (Canada) and the Nearctic Region. Aquatic Insects. [first publd online (11.i.2019)]

Orel, O. V., Semenchenko, A. A. (2019) Morphological description and DNA barcodes of adult males of Tanytarsus heliomesonyctios Langton, 1999 (Diptera, Chironomidae) in northeast of Russia. Zootaxa 4686(1): 119–126. [publd 14.x.2019]

Pinho, L. C., Dantas, G. P. S., Hamada, N. (2019) Some notes on taxonomy and distribution of Neotropical Aedokritus Roback and Axarus Roback (Diptera: Chironomidae: Chironominae). Zootaxa 4668(4): 535-542. [publd 12.ix.2019]

Silva, F. L. da (2019) Procladius travassosi (Oliveira, Messias & Silva-Vasconcelos, 1992) regarded as a new junior synonym of Procladius stroudi Roback 1982 (Diptera: Chironomidae: Tanypodinae). Zootaxa 4555(2): 280-282.

Silva, F. L. da, Stur, E. (2019) Pentaneurella katterjokki Fittkau & Murray (Chironomidae, Tanypodinae): redescription and phylogenetic position. ZooKeys 833: 107-119.

Silva, F. L.[Fabíola Lima] da, Dantas, G. P. S., Hamada, N. (2019) Description of immature stages of Ablabesmyia cordeiroi Neubern, 2013 (Diptera: Chironomidae: Tanypodinae). Acta Amazonica 49(2): 118-121.

Stur, E., Silva, F. L. da, Ekrem, T. (2019) Back from the past: DNA barcodes and morphology support Ablabesmyia americana Fittkau as a valid species (Diptera: Chironomidae). Diversity 11(9, paper 173): 15 pp. [publd online 19.ix.2019]

Wang, Q., Yu, H.-J., Wang, X.-H., Lin, X.-L. (2019) A newly recorded species, Sergentia kizakiensis (Tokunaga, 1940) (Diptera: Chironomidae), from Oriental China with DNA barcode. CHIRONOMUS J. Chironomidae Res. 32: 75-78. [publd online (19.xii.2019)]

Yamamoto, N., Suzuki, M., Yamamoto, M. (2019) Taxonomic notes on several Japanese chironomids (Diptera) described by Dr. M. Sasa(†) and his coauthors. Japan. J. syst. Ent. 25(1): 63-72. [publd]
Recently published / Re: New fossil taxa - all subf...
Last post by Martin Spies - December 16, 2019, 04:15:58 PM

Ashe, P., O'Connor, J. P. (2019) Validation of the fossil genus name †Acutiforcipia (Diptera: Chironomidae: Orthocladiinae). Bull. Irish Biogeogr. Soc. 43: 131-132.

Baranov, V., Hoffeins, C., Hoffeins, H.-W., Haug, J. T. (2019) Reaching across the ocean of time: A midge morphotype from the Cretaceous of Gondwana found in the Eocene Baltic amber. Palaeontologia Electronica 22.2.38A: 1-17. [][/color]

Giłka, W., Makarchenko, E. A., Pankowski, M. K., Zakrzewska, M. (2019) Myanmaro primus gen. et sp. nov., the first orthoclad (Diptera: Chironomidae) from Cretaceous Burmese amber. Zootaxa 4565(1): 61-70.

Giłka, W., Zakrzewska, M., Makarchenko, E. A. (2019) Burmochlus gen. nov., the first Cretaceous member of the Archaeochlus cluster (Diptera: Chironomidae: Podonominae). Cretaceous Res. 106: 8 pp. [first publd online (23.ix.2019)]

Krzemiński, W., Blagoderov, V., Azar, D., Lukashevich, E., Szadziewski, R., Wedmann, S., Nel, A., Collomb, F.-M., Waller, A., Nicholson, D. B. (2019) True flies (Insecta: Diptera) from the late Eocene insect limestone (Bembridge Marls) of the Isle of Wight, England, UK. Earth envir. Sci. Trans. R. Soc. Edinburgh 110(3-4, Sept.): 495-554.

Lukashevich, E. D., Przhiboro, A. A. (2019) A new genus of Chironomidae (Insecta: Diptera) from the Lower Cretaceous of Mongolia. Paleontol. J. 52(12): 1401-1407. ["2018" but publd (in print) 16.i.2019]
Identification / New publications
Last post by Martin Spies - December 09, 2019, 02:06:32 PM

Milošević, D., Milosavljević, A., Predić, B., Medeiros, A. S., Savić-Zdravković, D., Stojković Piperac, M., Kostić, T., Spasić, F., Leese, F. (2019) Application of deep learning in aquatic bioassessment: Towards automated identification of non-biting midges. Sci. total Envir.: 7 pp. + online suppl. material. [first publd online 14.xi.2019]
Identification / Remarks
Last post by Martin Spies - December 09, 2019, 02:03:49 PM
This board is intended for references or discussion concerning methods in chironomid identification.